Ecological succession refers to the process of change in the species composition and structure of an ecological community over time. There are two main theories that explain the process of ecological succession: the classical theory of succession and the modern theory of succession.
1. Classical Theory of Succession: The classical theory of succession, also known as the Clementsian theory, was proposed by Frederic Clements in the early 20th century. According to this theory, ecological succession is a predictable and deterministic process that follows a specific sequence of stages. It suggests that communities develop and progress towards a stable climax community, which is the final and most diverse stage of succession.
The classical theory emphasizes the role of biotic interactions, such as competition and facilitation, in driving the succession process. It suggests that each stage of succession prepares the environment for the next stage, leading to a gradual replacement of species until the climax community is reached. This theory assumes that the environment remains relatively stable during succession and that the climax community represents the optimal state for that particular ecosystem.
2. Modern Theory of Succession: The modern theory of succession, also known as the Gleasonian theory, was proposed by Henry Gleason in the 1920s as a critique of the classical theory. This theory argues that ecological succession is a more stochastic and unpredictable process, influenced by a combination of biotic and abiotic factors. It suggests that communities do not necessarily progress towards a stable climax community, but rather reach a dynamic equilibrium influenced by disturbances and environmental variability.
The modern theory emphasizes the role of chance events, such as disturbances (e.g., fire, storms) and dispersal limitations, in shaping the trajectory of succession. It suggests that communities are not predetermined but are assembled based on the availability of species and their ability to colonize and persist in a particular environment. This theory recognizes that succession can be reversible, with communities shifting back and forth between different stages depending on the prevailing conditions.
In summary, the classical theory of succession proposes a deterministic and predictable process leading to a stable climax community, while the modern theory suggests a more stochastic and dynamic process influenced by chance events and environmental variability. Both theories contribute to our understanding of ecological succession, and the debate between them continues to shape the field of ecology.